MPs must review a proposal to close 17 Scottish HMRC offices - including one in Glenrothes.
The government tax-collecting agency plans to move all its staff into 2 regional centres in Glasgow and Edinburgh. They say it will cut costs and save taxpayers money.
PCS general secretary Mark Serwotka said: "No one should be in any doubt that, if implemented, these proposals would be absolutely devastating for HMRC and the people who work there.
"Closing this many offices would pose a significant threat to the operation of HMRC, its service to the public and the working lives of staff, and the need for parliamentary scrutiny of the plans is undeniable and urgent."
Lin Homer, HMRC’s Chief Executive, said: "HMRC is committed to modern, regional centres serving every region and nation in the UK, with skilled and varied jobs and development opportunities, while also ensuring jobs are spread throughout the UK and not concentrated in the capital.
"HMRC has too many expensive, isolated and outdated offices. This makes it difficult for us to collaborate, modernise our ways of working, and make the changes we need to transform our service to customers and clamp down further on the minority who try to cheat the system.
"The new regional centres in Glasgow and Edinburgh will bring our staff together in more modern and cost-effective buildings in areas with lower rents. They will also make a big contribution to the Scottish economy, providing high-quality, skilled jobs and supporting the Government’s commitment to a national recovery that benefits all parts of the UK."
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